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In both years, borage inflorescences had the less
saponin content compared to its leaves (Table 5). The saponin content of
inflorescences in the second year was significantly lower than the first year.
Changes in saponin content of borage inflorescences in response to irrigation
intervals and seed priming were different in two years (Table 5). In the first
year, the saponin content of plants irrigated with 120 mm (I3) and
150 mm evaporation intervals (I4) was significantly higher than I1
and I2 treatments. Under moderate (I3) and severe stress
(I4), seed treatment with KNO3 and KH2PO4
significantly increased the saponin content of inflorescences. However in 2013,
saponin concentration of plants from primed and unprimed seeds was similar
under the different irrigation treatments (Table 5).

3.2.2. Tannin compounds content

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The tannin content of borage leaf and inflorescences
was statistically different in two years. Effects of irrigation and seed
priming and the interaction of these factors were not significant for tannin
content of leaf and inflorescences. However, leaf tannin content was also
significantly influenced by interactions of year × irrigation and year × seed
priming (Table 3).

The tannin content of borage
leaves in the second year was less than the first year at all irrigation levels
and different seed priming treatments. In the first year, there was no
significant difference between different irrigation intervals in leaf tannin
content. While in the second year, the lowest tannin concentration of leaves
was obtained under mild water stress (I2) and the leaf tannin
content of plants irrigated with 120 mm and 150 mm evaporation intervals were
12.5 and 5.3% higher than that plants irrigated with 60 mm evaporation
intervals (Fig. 3A). On the other hand in the 2012, there was no significant
difference between the plants from primed and unprimed seeds in terms of tannin
content of leaves, but in the 2013, seed priming led to increase the
concentration of tannin in the borage leaves (Fig. 3B). The concentration of
tannin compounds in borage leaves was more than that in inflorescences in both
years. Similar to leaf tannin content, tannin compounds content of
inflorescences in the second year was lower than that in the first year (Table
4).

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